COVER In Through the Looking-Glass and What Alice Found There, the Red Queen complained that in Wonderland "it takes all the running you can do, to keep in the same place." She has thus become the namesake of coevolutionary arms races in which plants continually adapt to retain resistance against their ever-evolving pathogens. The intricacies of the interactions between plants and their pathogens are explored in a special section beginning on page 2269. [David Galchutt]

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